Tupilakosaur-like vertebrae in Bothriceps australis, an Australian Brachyopid Stereospondyl

A new locality near the Permo-Triassic boundary in Tasmania, Australia, has yielded a number of specimens of the brachyopid stereospondyl, Bothriceps australis, whose provenance was unknown previously. An articulated diplospondylous vertebral column of a morphology typical only of the Tupilakosauridae was found at the same locality. Although the Brachyopidae and Tupilakosauridae have skulls of similar shape, most authors have regarded them as not closely related. Phylogenetic analysis confirms that the Brachyopidae and Tupilakosauridae are both members of the Stereospondyli but are not closely related, despite the remarkable similarity of the Bothriceps and tupilakosaur vertebral columns